Classification Of Fungi Alexopoulos And Mims 1979 Pdf File

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General trends and criteria for Algal classification Bold and Wynne, Ultrastructure of Prokaryotic and Eukaryotic algal cells and their components - cell wall, protoplasm, flagella, eye spots, chloroplast, pyrenoid, nucleus, pigments and reserve foods. Range of thallus structure, origin and evolution of sex in algae, phylogeny and interrelationships of algae.

Fungi Breakdown

Dikarya inc. Editor-In-Chief: C. Michael Gibson, M. The majority of species grow as multicellular filaments called hyphae forming a mycelium ; some fungal species also grow as single cells. Sexual and asexual reproduction of the fungi is commonly via spores , often produced on specialized structures or in fruiting bodies.

Some species have lost the ability to form specialized reproductive structures, and propagate solely by vegetative growth. Yeasts , molds , and mushrooms are examples of fungi. The fungi are a monophyletic group that is phylogenetically clearly distinct from the morphologically similar slime molds myxomycetes and water molds oomycetes. The fungi are more closely related to animals than plants , yet the discipline of biology devoted to the study of fungi, known as mycology , often falls under a branch of botany.

Occurring worldwide, most fungi are largely invisible to the naked eye, living for the most part in soil, dead matter, and as symbionts of plants, animals, or other fungi.

They perform an essential role in all ecosystems in decomposing organic matter and are indispensable in nutrient cycling and exchange. Some fungi become noticeable when fruiting, either as mushrooms or molds. Many fungal species have long been used as a direct source of food, such as mushrooms and truffles and in fermentation of various food products, such as wine , beer , and soy sauce. More recently, fungi are being used as sources for antibiotics used in medicine and various enzymes , such as cellulases , pectinases , and proteases , important for industrial use or as active ingredients of detergents.

Many fungi produce bioactive compounds called mycotoxins , such as alkaloids and polyketides that are toxic to animals including humans. Some fungi are used recreationally or in traditional ceremonies as a source of psychotropic compounds. Several species of the fungi are significant pathogens of humans and other animals, and losses due to diseases of crops e. The English word fungus is directly adopted from the Latin fungus , meaning "mushroom", used in Horace and Pliny. Fungi have a worldwide distribution, and grow in a wide range of habitats, including deserts.

Most fungi grow in terrestrial environments, but several species occur only in aquatic habitats. Fungi along with bacteria are the primary decomposers of organic matter in most if not all terrestrial ecosystems worldwide. Based on observations of the ratio of the number of fungal species to the number of plant species in some environments, the fungal kingdom has been estimated to contain about 1.

Human use of fungi for food preparation or preservation and other purposes is extensive and has a long history: yeasts are required for fermentation of beer , wine [9] and bread , some other fungal species are used in the production of soy sauce and tempeh. Mushroom farming and mushroom gathering are large industries in many countries. The study of the historical uses and sociological impact of fungi is known as ethnomycology.

Baker's yeast or Saccharomyces cerevisiae , a single-cell fungus, is used in the baking of bread and other wheat-based products, such as pizza and dumplings.

Fungi are also used extensively to produce industrial chemicals like lactic acid , antibiotics and even to make stonewashed jeans. Many fungi produce compounds with biological activity. Several of these compounds are toxic and are therefore called mycotoxins , referring to their fungal origin and toxic activity.

Of particular relevance to humans are those mycotoxins that are produced by moulds causing food spoilage and poisonous mushrooms see below. Particularly infamous are the aflatoxins , which are insidious liver toxins and highly carcinogenic metabolites produced by Aspergillus species often growing in or on grains and nuts consumed by humans, and the lethal amatoxins produced by mushrooms of the genus Amanita.

Other notable mycotoxins include ochratoxins , patulin , ergot alkaloids , and trichothecenes and fumonisins, all of which have significant impact on human food supplies or animal livestock.

Mycotoxins belong to the group of secondary metabolites or natural products. Originally, this group of compounds had been thought to be mere byproducts of primary metabolism , hence the name "secondary" metabolites. However, recent research has shown the existence of biochemical pathways solely for the purpose of producing mycotoxins and other natural products in fungi.

Some of the best known types of fungi are the edible and the poisonous mushrooms. Many species are commercially raised, but others must be harvested from the wild. Agaricus bisporus , sold as button mushrooms when small or Portobello mushrooms when larger, are the most commonly eaten species, used in salads, soups, and many other dishes.

Many Asian fungi are commercially grown and have gained in popularity in the West. They are often available fresh in grocery stores and markets, including straw mushrooms Volvariella volvacea , oyster mushrooms Pleurotus ostreatus , shiitakes Lentinula edodes , and enokitake Flammulina spp.

There are many more mushroom species that are harvested from the wild for personal consumption or commercial sale. Milk mushrooms , morels , chanterelles , truffles , black trumpets , and porcini mushrooms Boletus edulis also known as king boletes all demand a high price on the market.

They are often used in gourmet dishes. For certain types of cheeses , it is also a common practice to inoculate milk curds with fungal spores to foment the growth of specific species of mold that impart a unique flavor and texture to the cheese.

This accounts for the blue colour in cheeses such as Stilton or Roquefort which is created using Penicillium roqueforti spores. Many mushroom species are toxic to humans, with toxicities ranging from slight digestive problems or allergic reactions as well as hallucinations to severe organ failures and death.

Some of the most deadly mushrooms belong to the genera Inocybe , Cortinarius , and most infamously, Amanita. The latter genus includes the destroying angel A. Fly agaric mushrooms A. Historically Fly agaric was used by Celtic Druids in Northern Europe and the Koryak people of north-eastern Siberia for religious or shamanic purposes. In agricultural settings, fungi that actively compete for nutrients and space with, and eventually prevail over, pathogenic microorganisms, such as bacteria or other fungi, via the competitive exclusion principle , [25] or are parasites of these pathogens, may be beneficial agents for human use.

For example, some fungi may be used to suppress growth or eliminate harmful plant pathogens, such as insects, mites , weeds , nematodes and other fungi that cause diseases of important crop plants.

Entomopathogenic fungi can be used as biopesticides , as they actively kill insects. These alkaloids protect the infected grass plants from herbivory, but some endophyte alkaloids can cause poisoning of grazing animals, such as cattle and sheep. Certain fungi, in particular 'white rot' fungi, can degrade insecticides, herbicides, pentachlorophenol, creosote, coal tars, and heavy fuels and turn them into carbon dioxide, water, and basic elements.

Although often inconspicuous, fungi occur in every environment on Earth and play very important roles in most ecosystems. Along with bacteria, fungi are the major decomposers in most terrestrial and some aquatic ecosystems, and therefore play a critical role in biogeochemical cycles and in many food webs.

As decomposers, they play an indispensable role in nutrient cycling , especially as saprotrophs and symbionts , degrading organic matter to inorganic molecules, which can then re-enter anabolic metabolic pathways in plants or other organisms. Many fungi have important symbiotic relationships with organisms from most if not all Kingdoms.

In some mycorrhizal associations, the fungal partners may mediate plant-to-plant transfer of carbohydrates and other nutrients. Such mycorrhizal communities are called "common mycorrhizal networks". Lichens are formed by a symbiotic relationship between algae or cyanobacteria referred to in lichens as "photobionts" and fungi mostly various species of ascomycetes and a few basidiomycetes , in which individual photobiont cells are embedded in a tissue formed by the fungus.

The functions of both symbiotic organisms are so closely intertwined that they function almost as a single organism.

Many insects also engage in mutualistic relationships with various types of fungi. Several groups of ants cultivate fungi in the order Agaricales as their primary food source, while ambrosia beetles cultivate various species of fungi in the bark of trees that they infest. However, many fungi are parasites on plants, animals including humans , and other fungi.

Serious fungal pathogens of many cultivated plants causing extensive damage and losses to agriculture and forestry include the rice blast fungus Magnaporthe oryzae , [50] tree pathogens such as Ophiostoma ulmi and Ophiostoma novo-ulmi causing Dutch elm disease , [51] and Cryphonectria parasitica responsible for chestnut blight , [52] and plant-pathogenic fungi in the genera Fusarium , Ustilago , Alternaria , and Cochliobolus ; [40] fungi with the potential to cause serious human diseases, especially in persons with immuno-deficiencies , are in the genera Aspergillus , Candida , Cryptoccocus , [53] [41] [54] Histoplasma , [55] and Pneumocystis.

Some fungi are predators of nematodes , which they capture using an array of specialized structures, such as constricting rings or adhesive nets.

Growth of fungi as hyphae on or in solid substrates or single cells in aquatic environments is adapted to efficient extraction of nutrients from these environments, because these growth forms have high surface area to volume ratios. These adaptations in morphology are complemented by hydrolytic enzymes secreted into the environment for digestion of large organic molecules, such as polysaccharides , proteins , lipids , and other organic substrates into smaller molecules. Traditionally, the fungi are considered heterotrophs , organisms that rely solely on carbon fixed by other organisms for metabolism.

Fungi have evolved a remarkable metabolic versatility that allows many of them to use a large variety of organic substrates for growth, including simple compounds as nitrate , ammonia , acetate , or ethanol. Though fungi are part of the opisthokont clade, all phyla except for the chytrids have lost their posterior flagella. Many fungi grow as thread-like filamentous microscopic structures called hyphae , and an assemblage of intertwined and interconnected hyphae is called a mycelium.

However, septa have pores, such as the doliporus in the basidiomycetes that allow cytoplasm, organelles, and sometimes nuclei to pass through. Fungal mycelia can become visible macroscopically, for example, as concentric rings on various surfaces, such as damp walls, and on other substrates, such as spoilt food see figure , and are commonly and generically called mould American spelling , mold ; fungal mycelia grown on solid agar media in laboratory petri dishes are usually referred to as colonies, with many species exhibiting characteristic macroscopic growth morphologies and colours, due to spores or pigmentation.

Specialized fungal structures important in sexual reproduction are the apothecia , perithecia , and cleistothecia in the ascomycetes, and the fruiting bodies of the basidiomycetes, and a few ascomycetes.

These reproductive structures can sometimes grow very large, and are well known as mushrooms. Fungal hyphae are specifically adapted to growth on solid surfaces and within substrates, and can exert astoundingly large penetrative mechanical forces. The plant pathogen , Magnaporthe grisea , forms a structure called an appressorium specifically designed for penetration of plant tissues, and the pressure generated by the appressorium, which is directed against the plant epidermis can exceed 8 MPa 80 bars.

Reproduction of fungi is complex, reflecting the heterogeneity in lifestyles and genetic make up within this group of organisms. These conditions trigger genetically determined developmental programs leading to the expression of specialized structures for sexual or asexual reproduction.

These structures aid both reproduction and efficient dissemination of spores or spore-containing propagules. Asexual reproduction via vegetative spores or through mycelial fragmentation is common in many fungal species and allows more rapid dispersal than sexual reproduction.

In the case of the "Fungi imperfecti" or Deuteromycota , which lack a sexual cycle, it is the only means of propagation. Asexual spores, upon germination , may found a population that is clonal to the population from which the spore originated, and thus colonize new environments.

Sexual reproduction with meiosis exists in all fungal phyla, except the Deuteromycota. It differs in many aspects from sexual reproduction in animals or plants.

Many differences also exist between fungal groups and have been used to discriminate fungal clades and species based on morphological differences in sexual structures and reproductive strategies.

Experimental crosses between fungal isolates can also be used to identify species based on biological species concepts. The major fungal clades have initially been delineated based on the morphology of their sexual structures and spores; for example, the spore-containing structures, asci and basidia , can be used in the identification of ascomycetes and basidiomycetes, respectively. Many fungal species have elaborate vegetative incompatibility systems that allow mating only between individuals of opposite mating type , while others can mate and sexually reproduce with any other individual or itself.

Species of the former mating system are called heterothallic , and of the latter homothallic. Most fungi have both a haploid and diploid stage in their life cycles. In all sexually reproducing fungi, compatible individuals combine by cell fusion of vegetative hyphae by anastomosis , required for the initiation of the sexual cycle.

Ascomycetes and basidiomycetes go through a dikaryotic stage, in which the nuclei inherited from the two parents do not fuse immediately after cell fusion, but remain separate in the hyphal cells see heterokaryosis.

In ascomycetes, dikaryotic hyphae of the hymenium form a characteristic hook at the hyphal septum. During cell division formation of the hook ensures proper distribution of the newly divided nuclei into the apical and basal hyphal compartments. An ascus plural asci is then formed, in which karyogamy nuclear fusion occurs.

A General Framework of Persistence Strategies for Biological Systems Helps Explain Domains of Life

A fungus plural : fungi [2] or funguses [3] is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds , as well as the more familiar mushrooms. These organisms are classified as a kingdom , which is separate from the other eukaryotic life kingdoms of plants and animals. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs ; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize.

Revised and updated in accordance with modern taxonomic proposals, this edition offers a well--documented, logical and clear explanation of the structure and classification of fungi along with an introduction to physiological, biochemical, genetic and ecological data. The taxonomic approach provides a framework with predictive value. Read more HathiTrust Digital Library, Limited view search only. Please choose whether or not you want other users to be able to see on your profile that this library is a favorite of yours.

Introductory mycology

Reisen und Leben im Wohnmobil. Accueil Contact. Five kingdoms Alexopoulos and Mims,

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The nature and cause of the division of organisms in superkingdoms is not fully understood. Assuming that environment shapes physiology, here we construct a novel theoretical framework that helps identify general patterns of organism persistence. Scope denotes the entirety of environmental events and conditions to which the organism is exposed during its lifetime. These concepts bring organisms of different complexity to a common ecological denominator. Ecological and physiological data suggest organisms persist using three strategies: flexibility, robustness, and economy.

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Dikarya inc. Editor-In-Chief: C. Michael Gibson, M. The majority of species grow as multicellular filaments called hyphae forming a mycelium ; some fungal species also grow as single cells. Sexual and asexual reproduction of the fungi is commonly via spores , often produced on specialized structures or in fruiting bodies. Some species have lost the ability to form specialized reproductive structures, and propagate solely by vegetative growth.

One-stop Solution. Beta Lifescience. In this article we will discuss about the classification of fungi by various botanists. But the detailed classification of fungi was first proposed by Elias Read Next Story. Later, much more handy system of classification was proposed by Gwynne- Vaughan and Barnes In their classification, Myxomycetes were not Monocot and Dicot Roots With Diagram Plants considered as fungi and were included by them as forms resembling fungi.

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